ABOUT ELEPHANTS

Elephants are large land mammals of the order Proboscidea and the family Elephantidae. There are three living species: the African Bush Elephant, the African Forest Elephant and the Asian Elephant (also known as the Indian Elephant). Other species have become extinct since the last ice age, the Mammoths, dwarf forms of which may have survived as late as 2,000 BC, being the best-known of these. They were once classified along with other thick skinned animals in a now invalid order, Pachydermata. Elephants are the largest land animals. The elephant's gestation period is 22 months, the longest of any land animal. At birth it is common for an elephant calf to weigh 120 kilograms (260 lb). They typically live for 50 to 70 years, but the oldest recorded elephant lived for 82 years. The largest elephant ever recorded was shot in Angola in 1956. This male weighed about 12,000 kilograms (26,000 lb), with a shoulder height of 4.2 metres (14 ft), a metre (yard) taller than the average male African elephant. The smallest elephants, about the size of a calf or a large pig, were a prehistoric species that lived on the island of Crete during the Pleistocene epoch. The elephant has appeared in cultures across the world. They are a symbol of wisdom in Asian cultures and are famed for their memory and intelligence, where they are thought to be on par with cetaceans and hominids. Healthy adult elephants have no natural predators, although lions may take calves or weak individuals. They are, however, increasingly threatened by human intrusion and poaching. Once numbering in the millions, the African elephant population has dwindled to between 470,000 and 690,000 individuals according to a March 2007 estimate. While the elephant is a protected species worldwide, with restrictions in place on capture, domestic use, and trade in products such as ivory has resulted in increased poaching. Certain African nations report a decrease of their elephant populations by as much as two-thirds, and populations in certain protected areas are in danger of being eliminated. The African Elephant genus contains two (or, arguably, three) living species; whereas the Asian Elephant species is the only surviving member of the Asian Elephant genus, but can be divided into four subspecies.

Although the fossil evidence is uncertain, scientists discovered genetic evidence that the elephant family shares distant ancestry with the sirenians (sea cows) and the hyraxes through gene comparisons. In the distant past, members of the hyrax family grew to large sizes, and it seems likely that the common ancestor of all three modern families was some kind of amphibious hyracoid. One theory suggests that these animals spent most of their time under water, using their trunks like snorkels for breathing. Modern elephants have retained this ability and are known to swim in that manner for up to 6 hours and 50 km (30 miles). In the past, there was a much wider variety of elephant genera, including the mammoths and stegodons. There was also a much wider variety of species.

The Elephants of the genus Loxodonta, known collectively as African elephants, are currently found in 37 countries in Africa. African elephants are distinguished from Asian elephants in several ways, the most noticeable being their ears which are much larger. The African elephant is typically larger than the Asian elephant and has a concave back. Both African males and females have external tusks and are usually less hairy than their Asian cousins. African elephants have traditionally been classified as a single species comprising two distinct subspecies, namely the savanna elephant (Loxodonta africana africana) and the forest elephant (Loxodonta africana cyclotis), but recent DNA analysis suggests that these may actually constitute distinct species. This split is not universally accepted by experts and a third species of African elephant has also been proposed. This reclassification has important implications for conservation, because it means that where previously it was assumed that a single and endangered species comprised two small populations, if in reality these are two separate species, then as a consequence, both could be more gravely endangered than a more numerous and wide-ranging single species might have been. There is also a potential danger in that, if the forest elephant is not explicitly listed as an endangered species, poachers and smugglers might be able to evade the law forbidding trade in endangered animals and their body parts. The Forest elephant and the Savanna elephant can also hybridise that is, breed together successfully, though their preferences for different terrains reduce such opportunities. As the African elephant has only recently been recognized to comprise two separate species, groups of captive elephants have not been comprehensively classified and some could well be hybrids. Under the new two species classification, Loxodonta africana refers specifically to the Savanna Elephant, the largest of all elephants. In fact, it is the largest land animal in the world, with the males standing 3.2 metres (10 ft) to 4 metres (13 ft) at the shoulder and weighing 3,500 kilograms (7,700 lb) to a reported 12,000 kilograms (26,000 lb). The female is smaller, standing about 3 metres (9.8 ft) at the shoulder. Most often, Savanna Elephants are found in open grasslands, marshes, and lakeshores. They range over much of the savanna zone south of the Sahara. The other putative species, the Forest Elephant (Loxodonta cyclotis), is usually smaller and rounder, and its tusks thinner and straighter compared with the Savanna Elephant. The Forest Elephant can weigh up to 4,500 kilograms (9,900 lb) and stand about 3 metres (10 ft) tall. Much less is known about these animals than their savanna cousins, because environmental and political obstacles make them difficult to study. Normally, they inhabit the dense African rain forests of central and western Africa, although occasionally they roam the edges of forests, thus overlapping the Savanna elephant territories and hybridizing.

Elephant populations in West Africa, on the other hand, are generally small and fragmented, and only account for a small proportion of the continental total. Much uncertainty remains as to the size of the elephant population in Central Africa, where the prevalence of forest makes population surveys difficult, but poaching for ivory and bushmeat is believed to be intense through much of the region.


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The Asian elephant, Elephas maximus, is smaller than the African. It has smaller ears, and typically, only the males have large external tusks. The world population of Asian elephants also called Indian Elephants is estimated to be around 60,000, about a tenth of the number of African elephants. More precisely, it is estimated that there are between 38,000 and 53,000 wild elephants and between 14,500 and 15,300 domesticated elephants in Asia with perhaps another 1,000 scattered around zoos in the rest of the world. The Asian elephants' decline has possibly been more gradual than the African and caused primarily by poaching and habitat destruction by human encroachment. Several subspecies of Elephas maximus have been identified, using morphometric data and molecular markers. Elephas maximus maximus (Sri Lankan Elephant) is found only on the island of Sri Lanka. It is the largest of the Asians. There are an estimated 3,000-4,500 members of this subspecies left today in the wild, although no accurate census has been carried out recently. Large males can weigh upward to 5,400 kg (12,000 lb) and stand over 3.4 m (11 ft) tall. Sri Lankan males have very large cranial bulges, and both sexes have more areas of depigmentation than other Asians. Typically, their ears, face, trunk, and belly have large concentrations of pink-speckled skin. There is an orphanage for elephants in Pinnawala, Sri Lanka, which plays a large role in protecting the Sri Lankan Elephant from extinction. Elephas maximus indicus (Indian Elephant) makes up the bulk of the Asian elephant population. Numbering approximately 36,000, these elephants are lighter grey in colour, with depigmentation only on the ears and trunk. Large males will ordinarily weigh only about 5,000 kg (11,000 lb), but are as tall as the Sri Lankan. The mainland Asian can be found in 11 Asian countries, from India to Indonesia. They prefer forested areas and transitional zones, between forests and grasslands, where greater food variety is available. The smallest of all the elephants is the Sumatran Elephant, Elephas maximus sumatranus. Population estimates for this group range from 2,100 to 3,000 individuals. It is very light grey in colour and has less depigmentation than the other Asians, with pink spots only on the ears. Mature Sumatrans will usually only measure 1.7-2.6 m (5.6-8.5 ft) at the shoulder and weigh less than 3,000 kg (6,600 lb). It is considerably smaller than its other Asian (and African) cousins and exists only on the island of Sumatra, usually in forested regions and partially wooded habitats. In 2003, a further subspecies was identified on Borneo. Named the Borneo pygmy elephant, it is smaller and tamer than any other Asian elephants. It also has relatively larger ears, longer tail and straighter tusks.

The proboscis, or trunk, is a fusion of the nose and upper lip, elongated and specialized to become the elephant's most important and versatile appendage. African elephants are equipped with two fingerlike projections at the tip of their trunk, while Asians have only one. According to biologists, the elephant's trunk may have over forty thousand individual muscles in it, making it sensitive enough to pick up a single blade of grass, yet strong enough to rip the branches off a tree. Most herbivores (plant eaters, like the elephant) possess teeth adapted for cutting and tearing off plant materials. However, except for the very young or infirm, elephants always use their trunks to tear up their food and then place it in their mouth. They will graze on grass or reach up into trees to grasp leaves, fruit, or entire branches. If the desired food item is too high up, the elephant will wrap its trunk around the tree or branch and shake its food loose or sometimes simply knock the tree down altogether. The trunk is also used for drinking. Elephants suck water up into the trunk (up to fifteen quarts or fourteen litres at a time) and then blow it into their mouth. Elephants also inhale water to spray on their body during bathing. On top of this watery coating, the animal will then spray dirt and mud, which act as a protective sunscreen. When swimming, the trunk makes an excellent snorkel. This appendage also plays a key role in many social interactions. Familiar elephants will greet each other by entwining their trunks, much like a handshake. They also use them while play-wrestling, caressing during courtship and mother / child interactions, and for dominance displays a raised trunk can be a warning or threat, while a lowered trunk can be a sign of submission. Elephants can defend themselves very well by flailing their trunk at unwanted intruders or by grasping and flinging them. An elephant also relies on its trunk for its highly developed sense of smell. Raising the trunk up in the air and swivelling it from side to side, like a periscope, it can determine the location of friends, enemies, and food sources.

The tusks of an elephant are its second upper incisors. Tusks grow continuously; an adult male's tusks will grow about 18 cm (7 in) a year. Tusks are used to dig for water, salt, and roots; to debark trees, to eat the bark; to dig into baobab trees to get at the pulp inside; and to move trees and branches when clearing a path. In addition, they are used for marking trees to establish territory and occasionally as weapons. Like humans who are typically right- or left-handed, elephants are usually right- or left-tusked. The dominant tusk, called the master tusk, is generally shorter and more rounded at the tip from wear. Both male and female African elephants have large tusks that can reach over 3 m (10 ft) in length and weigh over 90 kg (200 lb). In the Asian species, only the males have large tusks. Female Asians have tusks which are very small or absent altogether. Asian males can have tusks as long as the much larger Africans, but they are usually much slimmer and lighter; the heaviest recorded is 39 kg (86 lb). The tusk of both species is mostly made of calcium phosphate in the form of apatite. As a piece of living tissue, it is relatively soft (compared with other minerals such as rock), and the tusk, also known as ivory, is strongly favoured by artists for its carvability. The desire for elephant ivory has been one of the major factors in the reduction of the world's elephant population.

Elephants' teeth are very different from those of most other mammals. Over their lives they usually have 28 teeth.

Elephants are colloquially called pachyderms (from their original scientific classification), which means thick-skinned animals. An elephant's skin is extremely tough around most parts of its body and measures about 2.5 centimetres (1.0 in) thick. However, the skin around the mouth and inside of the ear is paper-thin. Normally, the skin of an Asian is covered with more hair than its African counterpart. This is most noticeable in the young. Asian calves are usually covered with a thick coat of brownish red fuzz. As they get older, this hair darkens and becomes more sparse, but it will always remain on their heads and tails. The species of elephants are typically greyish in colour, but the Africans very often appear brown or reddish from wallowing in mud holes of coloured soil. Wallowing is an important behaviour in elephant society. Not only is it important for socialization, but the mud acts as a sunscreen, protecting their skin from harsh ultraviolet radiation. Although tough, an elephant's skin is very sensitive. Without regular mud baths to protect it from burning, as well as from insect bites and moisture loss, an elephant's skin would suffer serious damage. After bathing, the elephant will usually use its trunk to blow dirt on its body to help dry and bake on its new protective coat. As elephants are limited to smaller and smaller areas, there is less water available, and local herds will often come too close over the right to use these limited resources. Wallowing also aids the skin in regulating body temperatures. Elephants have difficulty in releasing heat through the skin because, in proportion to their body size, they have very little of it. The ratio of an elephant's mass to the surface area of its skin is many times that of a human. Elephants have even been observed lifting up their legs to expose the soles of their feet, presumably in an effort to expose more skin to the air. Since wild elephants live in very hot climates, they must have other means of getting rid of excess heat.

An elephant's legs are great straight pillars, as they must be to support its bulk. The elephant needs less muscular power to stand because of its straight legs and large pad-like feet. For this reason an elephant can stand for very long periods of time without tiring. In fact, African elephants rarely lie down unless they are sick or wounded. Indian elephants, in contrast, lie down frequently. The feet of an elephant are nearly round. African elephants have three nails on each hind foot, and four on each front foot. Indian elephants have four nails on each hind foot and five on each front foot. Beneath the bones of the foot is a tough, gelatinous material that acts as a cushion or shock absorber. Under the elephant's weight the foot swells, but it gets smaller when the weight is removed. Elephants swim well, but cannot trot, jump, or gallop. They do have two gaits: a walk; and a faster gait that is similar to running. In walking, the legs act as pendulums, with the hips and shoulders rising and falling while the foot is planted on the ground. With no "aerial phase", the faster gait does not meet all the criteria of running, as elephants always have at least one foot on the ground. However, an elephant moving fast uses its legs much like a running animal, with the hips and shoulders falling and then rising while the feet are on the ground. In this gait, an elephant will have three feet off the ground at one time. As both of the hind feet and both of the front feet are off the ground at the same time, this gait has been likened to the hind legs and the front legs taking turns running. Although they start this "run" at only 8 km/h, elephants can reach speeds up to 40 km/h (25 mph), all the while using the same gait. At this speed, most other four-legged creatures are well into a gallop, even accounting for leg length. Spring-like kinetics could explain the difference between the motion of elephants and other animals.

The large flapping ears of an elephant are also very important for temperature regulation. Elephant ears are made of a very thin layer of skin stretched over cartilage and a rich network of blood vessels. On hot days, elephants will flap their ears constantly, creating a slight breeze. This breeze cools the surface blood vessels, and then the cooler blood gets circulated to the rest of the animal's body. The hot blood entering the ears can be cooled as much as ten degrees Fahrenheit before returning to the body. Differences in the ear sizes of African and Asian elephants can be explained, in part, by their geographical distribution. Africans originated and stayed near the equator, where it is warmer. Therefore, they have bigger ears. Asians live farther north, in slightly cooler climates, and thus have smaller ears. The ears are also used in certain displays of aggression and during the males' mating period. If an elephant wants to intimidate a predator or rival, it will spread its ears out wide to make itself look more massive and imposing. During the breeding season, males give off an odour from the musth gland located behind their eyes.

Elephants live in a structured social order. The social lives of male and female elephants are very different. The females spend their entire lives in tightly knit family groups made up of mothers, daughters, sisters, and aunts. These groups are led by the eldest female, or matriarch. Adult males, on the other hand, live mostly solitary lives. The social circle of the female elephant does not end with the small family unit. In addition to encountering the local males that live on the fringes of one or more groups, the female's life also involves interaction with other families, clans, and subpopulations. Most immediate family groups range from five to fifteen adults, as well as a number of immature males and females. When a group gets too big, a few of the elder daughters will break off and form their own small group. They remain very aware of which local herds are relatives and which are not. The life of the adult male is very different. As he gets older, he begins to spend more time at the edge of the herd, gradually going off on his own for hours or days at a time. Eventually, days become weeks, and somewhere around the age of fourteen, the mature male, or bull, sets out from his natal group for good. While males do live primarily solitary lives, they will occasionally form loose associations with other males. These groups are called bachelor herds. The males spend much more time than the females fighting for dominance with each other. Only the most dominant males will be permitted to breed with cycling females. The less dominant ones must wait their turn. It is usually the older bulls, forty to fifty years old, that do most of the breeding. The dominance battles between males can look very fierce, but typically they inflict very little injury. Most of the bouts are in the form of aggressive displays and bluffs. Ordinarily, the smaller, younger, and less confident animal will back off before any real damage can be done. However, during the breeding season, the battles can get extremely aggressive, and the occasional elephant is injured. During this season, known as musth, a bull will fight with almost any other male it encounters, and it will spend most of its time hovering around the female herds, trying to find a receptive mate.

Elephants have well innervated trunks, and an exceptional sense of hearing and smell. The hearing receptors reside not only in ears, but also in trunks that are sensitive to vibrations, and most significantly feet, which have special receptors for low frequency sound and are exceptionally well innervated. Elephants communicate by sound over large distances of several kilometers partly through the ground, which is important for their social lives. Elephants are observed listening by putting trunks on the ground and carefully positioning their feet.

In addition to their bellows, roars and widely recognized trumpet-like calls; elephants communicate over long distances by producing and receiving low-frequency sound (infrasound), a sub-sonic rumbling, which can travel through the ground farther than sound travels through the air. This can be felt by the sensitive skin of an elephant's feet and trunk, which pick up the resonant vibrations much as the flat skin on the head of a drum.

Elephants are herbivores, spending 16 hours a day collecting plant food. Their diet is at least 50% grasses, supplemented with leaves, bamboo, twigs, bark, roots, and small amounts of fruits, seeds and flowers. Because elephants only digest 40% of what they eat, they have to make up for their digestive system's lack of efficiency in volume. An adult elephant can consume 140-270 kg (300-600 lb) of food a day.

Elephant social life revolves around breeding and raising of the calves. A female will usually be ready to breed around the age of thirteen, when she comes into estrus, a short phase of receptiveness lasting a couple of days, for the first time. Females announce their estrus with smell signals and special calls.

Females prefer bigger, stronger, and, most importantly, older males. Such a reproductive strategy tends to increase their offspring's chances of survival. After a twenty-two-month pregnancy, the mother will give birth to a calf that will weigh about 113 kg (250 lb) and stand over 76 cm (2.5 ft) tall. Elephants have a very long childhood. They are born with fewer survival instincts than many other animals. Instead, they must rely on their elders to teach them the things they need to know. Today, however, the pressures humans have put on the wild elephant populations, from poaching to habitat destruction, mean that the elderly often die at a younger age, leaving fewer teachers for the young. A new calf is usually the centre of attention for all herd members. All the adults and most of the other young will gather around the newborn, touching and caressing it with their trunks. The baby is born nearly blind and at first relies, almost completely, on its trunk to discover the world around it. As everyone in the herd is usually related, all members of the tightly knit female group participate in the care and protection of the young. After the initial excitement, the mother will usually select several full-time baby-sitters, or "allomothers", from her group.

Elephants are a species which many other organisms depend on. One particular example of that are termites mounds: termites eat elephant feces and often begin building their mounds under piles of elephant feces. Elephants' foraging activities can sometimes greatly affect the areas in which they live. By pulling down trees to eat leaves, breaking branches, and pulling out roots they create clearings in which new young trees and other vegetation can establish itself. During the dry season, elephants use their tusks to dig into dry river beds to reach underground sources of water. These newly dug water holes may then become the only source of water in the area. Elephants make pathways through their environment which are also used by other animals to access areas normally out of reach. These pathways have sometimes been used by several generations of elephants and today are converted by humans to paved roads.

War elephants were used by armies in the Indian sub-continent, the Warring States of China, and later by the Persian Empire. This use was adopted by Hellenistic armies after Alexander the Great experienced their worth against king Porus, notably in the Ptolemaic and Seleucid diadoch empires. The Carthaginian general Hannibal took elephants across the Alps when he was fighting the Romans, but brought too few elephants to be of much military use, although his horse cavalry was quite successful; he probably used a now-extinct third African (sub)species, the North African (Forest) elephant, smaller than its two southern cousins, and presumably easier to domesticate. A large elephant in full charge could cause tremendous damage to infantry, and cavalry horses would be afraid of them.


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