Charles Darwin (12 February 1809 - 19 April 1882) was an English naturalist, eminent as a collector and geologist, who proposed and provided scientific evidence that all species of life have evolved over time from common ancestors through the process he called natural selection. The fact that evolution occurs became accepted by the scientific community and the general public in his lifetime, while his theory of natural selection came to be widely seen as the primary explanation of the process of evolution in the 1930s, and now forms the basis of modern evolutionary theory. In modified form, Darwin’s scientific discovery remains the foundation of biology, as it provides a unifying logical explanation for the diversity of life. Charles Darwin developed his interest in natural history while studying first medicine at Edinburgh University, then theology at Cambridge. His five-year voyage on the Beagle established him as a geologist whose observations and theories supported Charles Lyell’s uniformitarian ideas, and publication of his journal of the voyage made him famous as a popular author. Puzzled by the geographical distribution of wildlife and fossils he collected on the voyage, Charles Darwin investigated the transmutation of species and conceived his theory of natural selection in 1838. Although he discussed his ideas with several naturalists, he needed time for extensive research and his geological work had priority. Charles Darwin was writing up his theory in 1858 when Alfred Russel Wallace sent him an essay which described a similar theory, prompting immediate joint publication of both of their theories. Charles Darwin 1859 book On the Origin of Species established evolution by common descent as the dominant scientific explanation of diversification in nature. He examined human evolution and sexual selection in The Descent of Man, and Selection in Relation to Sex, followed by The Expression of the Emotions in Man and Animals. Charles Darwin research on plants was published in a series of books, and in his final book, he examined earthworms and their effect on soil. In recognition of Darwin’s pre-eminence, he was buried in Westminster Abbey, close to John Herschel and Isaac Newton.

Charles Robert Darwin was born in Shrewsbury, Shropshire, England on 12 February 1809 at his family home, the Mount. Charles Darwin was the fifth of six children of wealthy society doctor and financier Robert Darwin, and Susannah Darwin (née Wedgwood). Charles Darwin was the grandson of Erasmus Darwin on his father’s side, and of Josiah Wedgwood on his mother’s side. Both families were largely Unitarian, though the Wedgwoods were adopting Anglicanism. Robert Darwin, himself quietly a freethinker, made a nod toward convention by having baby Charles baptised in the Anglican Church. Nonetheless, Charles and his siblings attended the Unitarian chapel with their mother, and in 1817, Charles joined the day school, run by its preacher. In July of that year, when Charles was eight years old, his mother died. From September 1818, he attended the nearby Anglican Shrewsbury School as a boarder. Charles Darwin spent the summer of 1825 as an apprentice doctor, helping his father treat the poor of Shropshire. In the autumn, he went to the University of Edinburgh, Scotland, to study medicine, but he was revolted by the brutality of surgery and neglected his medical studies. Charles Darwin learned taxidermy from John Edmonstone, a freed black slave who told him exciting tales of the South American rainforest. Later, in The Descent of Man, he used this experience as evidence that Negroes and Europeans were closely related despite superficial differences in appearance. In Darwin’s second year, he joined the Plinian Society, a student group interested in natural history. He became a keen pupil of Robert Edmund Grant, a proponent of Jean-Baptiste Lamarck’s theory of evolution by acquired characteristics, which Charles’s grandfather Erasmus had also advocated. On the shores of the Firth of Forth, Charles Darwin joined in Grant’s investigations of the life cycle of marine animals. These studies found evidence for homology, the radical theory that all animals have similar organs which differ only in complexity, thus showing common descent. In March 1827, Darwin made a presentation to the Plinian of his own discovery that the black spores often found in oyster shells were the eggs of a skate leech. Charles Darwin also sat in on Robert Jameson’s natural history course, learning about stratigraphic geology, receiving training in classifying plants, and assisting with work on the extensive collections of the University Museum, one of the largest museums in Europe at the time.

In 1827, his father, unhappy at his younger son’s lack of progress, shrewdly enrolled him in a Bachelor of Arts course at Christ’s College, Cambridge to qualify as a clergyman, expecting him to get a good income as an Anglican parson. However, Charles Darwin preferred riding and shooting to studying. Along with his cousin William Darwin Fox, Charles Darwin became engrossed in the craze at the time for the competitive collecting of beetles. Fox introduced him to the Reverend John Stevens Henslow, professor of botany, for expert advice on beetles. Darwin subsequently joined Henslow’s natural history course and became his favourite pupil, known to the dons as the man who walks with Henslow. When exams drew near, Darwin focused on his studies and received private instruction from Henslow. Charles Darwin was particularly enthusiastic about the writings of William Paley, including the argument for divine design in nature. It has been argued that Darwin’s enthusiasm for Paley’s religious adaptationism paradoxically played a role even later, when Darwin formulated his theory of natural selection. In his finals in January 1831, Charles Darwin performed well in theology and, having scraped through in classics, mathematics and physics, came tenth out of a pass list of 178. Residential requirements kept Darwin at Cambridge until June. Following Henslow’s example and advice, he was in no rush to take Holy Orders. Inspired by Alexander von Humboldt’s Personal Narrative, he planned to visit Tenerife with some classmates after graduation to study natural history in the tropics. To prepare himself, Charles Darwin joined the geology course of the Reverend Adam Sedgwick and, in the summer, went with him to assist in mapping strata in Wales. After a fortnight with student friends at Barmouth, he returned home to find a letter from Henslow recommending Darwin as a suitable (if unfinished) naturalist for the unpaid position of gentleman’s companion to Robert FitzRoy, the captain of HMS Beagle, which was to leave in four weeks on an expedition to chart the coastline of South America. His father objected to the planned two-year voyage, regarding it as a waste of time, but was persuaded by his brother-in-law, Josiah Wedgwood, to agree to his son’s participation.

The Beagle survey took five years, two-thirds of which Charles Darwin spent on land. He carefully noted a rich variety of geological features, fossils and living organisms, and methodically collected an enormous number of specimens, many of them new to science. At intervals during the voyage he sent specimens to Cambridge together with letters about his findings, and these established his reputation as a naturalist. His extensive detailed notes showed his gift for theorising and formed the basis for his later work. The journal he originally wrote for his family, published as The Voyage of the Beagle, summarises his findings and provides social, political and anthropological insights into the wide range of people he met, both native and colonial. While on board the ship, Darwin suffered badly from seasickness. In October 1833, Charles Darwin caught a fever in Argentina, and in July 1834, while returning from the Andes down to Valparaíso, he fell ill and spent a month in bed. Before they set out, FitzRoy gave Darwin the first volume of Charles Lyell’s Principles of Geology, which explained landforms as the outcome of gradual processes over huge periods of time. On their first stop ashore at St Jago, Darwin found that a white band high in the volcanic rock cliffs consisted of baked coral fragments and shells. This matched Lyell’s concept of land slowly rising or falling, giving Charles Darwin a new insight into the geological history of the island which inspired him to think of writing a book on geology. Charles Darwin went on to make many more discoveries, some of them particularly dramatic. Charles Darwin saw stepped plains of shingle and seashells in Patagonia as raised beaches, and after experiencing an earthquake in Chile saw mussel-beds stranded above high tide showing that the land had just been raised. High in the Andes he saw several fossil trees that had grown on a sand beach, with seashells nearby. Charles Darwin theorised that coral atolls form on sinking volcanic mountains, and confirmed this when the Beagle surveyed the Cocos (Keeling) Islands. In South America, Charles Darwin found and excavated rare fossils of gigantic extinct mammals in strata with modern seashells, indicating recent extinction and no change in climate or signs of catastrophe. Though he correctly identified one as a Megatherium and fragments of armour reminded him of the local armadillo, he assumed his finds were related to African or European species and it was a revelation to him after the voyage when Richard Owen showed that they were closely related to living creatures exclusively found in the Americas.

As HMS Beagle surveyed the coasts of South America, Charles Darwin began to theorise about the wonders of nature around him.Lyell’s second volume, which argued against evolutionism and explained species distribution by centres of creation, was sent out to Darwin. Charles Darwin puzzled over all he saw, and his ideas went beyond Lyell. In Argentina, he found that two types of rhea had separate but overlapping territories. On the Galápagos Islands he collected birds, and noted that mockingbirds differed depending on which island they came from. Charles Darwin also heard that local Spaniards could tell from their appearance on which island tortoises originated, but thought the creatures had been imported by buccaneers. In Australia, the marsupial rat-kangaroo and the platypus seemed so unusual that Charles Darwin thought it was almost as though two distinct Creators had been at work. In Cape Town he and FitzRoy met John Herschel, who had recently written to Lyell about that mystery of mysteries, the origin of species. When organising his notes on the return journey, Darwin wrote that if his growing suspicions about the mockingbirds, the tortoises and the Falkland Island Fox were correct, such facts undermine the stability of Species, then cautiously added would before undermine. Charles Darwin later wrote that such facts seemed to me to throw some light on the origin of species. Three natives who had been taken from Tierra del Fuego on the Beagle’s previous voyage were taken back there to become missionaries. They had become civilised in England over the previous two years, yet their relatives appeared to Charles Darwin to be miserable, degraded savages. A year on, the mission had been abandoned and only Jemmy Button spoke with them to say he preferred his harsh previous way of life and did not want to return to England. Because of this experience, Charles Darwin came to think that humans were not as far removed from animals as his friends then believed, and saw differences as relating to cultural advances towards civilisation rather than being racial. Captain FitzRoy was committed to writing the official Narrative of the Beagle voyages, and near the end of the voyage, he read Darwin’s diary and asked him to rewrite this Journal to provide the third volume, on natural history.

While Charles Darwin was still on the voyage, Henslow fostered his former pupil’s reputation by giving selected naturalists access to the fossil specimens and a pamphlet of Darwin’s geological letters. When the Beagle returned on 2 October 1836, Charles Darwin was a celebrity in scientific circles. After visiting his home in Shrewsbury and seeing relatives, Charles Darwin hurried to Cambridge to see Henslow, who advised on finding naturalists available to describe and catalogue the collections, and agreed to take on the botanical specimens. Darwin’s father organised investments, enabling his son to be a self-funded gentleman scientist, and an excited Darwin went round the London institutions being fêted and seeking experts to describe the collections. Zoologists had a huge backlog of work, and there was a danger of specimens just being left in storage. An eager Charles Lyell met Darwin for the first time on 29 October and soon introduced him to the up-and-coming anatomist Richard Owen, who had the facilities of the Royal College of Surgeons at his disposal to work on the fossil bones collected by Darwin. Owen’s surprising results included gigantic sloths, a hippopotamus-like skull from the extinct rodent Toxodon, and armour fragments from a huge extinct armadillo (Glyptodon), as Darwin had initially surmised. The fossil creatures were unrelated to African animals, but closely related to living species in South America. In mid-December, Darwin moved to Cambridge to organise work on his collections and rewrite his Journal. Charles Darwin wrote his first paper, showing that the South American landmass was slowly rising, and with Lyell’s enthusiastic backing read it to the Geological Society of London on 4 January 1837. On the same day, Charles Darwin presented his mammal and bird specimens to the Zoological Society. The ornithologist John Gould soon revealed that the Galapagos birds that Darwin had thought a mixture of blackbirds, gros-beaks and finches, were, in fact, twelve separate species of finches. On 17 February 1837, Charles Darwin was elected to the Council of the Geographical Society, and in his presidential address, Lyell presented Owen’s findings on Darwin’s fossils, stressing geographical continuity of species as supporting his uniformitarian ideas.

On 6 March 1837, Charles Darwin moved to London to be close to this work, and joined the social whirl around scientists and savants such as Charles Babbage, who thought that God preordained life by natural laws rather than ad hoc miraculous creations. Charles Darwin lived near his freethinking brother Erasmus, who was part of this Whig circle and whose close friend the writer Harriet Martineau promoted the ideas of Thomas Malthus underlying the Whig Poor Law reforms aimed at discouraging the poor from breeding beyond available food supplies. John Herschel’s question on the origin of species was widely discussed. Medical men even joined Grant in endorsing transmutation of species, but to Charles Darwin’s scientist friends such radical heresy attacked the divine basis of the social order already under threat from recession and riots. Gould now revealed that the Galapagos mockingbirds from different islands were separate species, not just varieties, and the wrens were yet another species of finches. Charles Darwin had not kept track of which islands the finch specimens were from, but found information from the notes of others on the Beagle, including FitzRoy, who had more carefully recorded their own collections. The zoologist Thomas Bell showed that the Galápagos tortoises were native to the islands. By mid-March, Darwin was convinced that creatures arriving in the islands had become altered in some way to form new species on the different islands, and investigated transmutation while noting his speculations in his Red Notebook which he had begun on the Beagle. In mid-July, he began his secret B notebook on transmutation, and on page 36 wrote I think above his first sketch of an evolutionary tree.

As well as launching into this intensive study of transmutation, Charles Darwin became mired in more work. While still rewriting his Journal, he took on editing and publishing the expert reports on his collections, and with Henslow’s help obtained a Treasury grant of £1,000 to sponsor this multi-volume Zoology of the Voyage of H.M.S. Beagle. Charles Darwin agreed to unrealistic dates for this and for a book on South American Geology supporting Lyell’s ideas. Charles Darwin finished writing his Journal around 20 June 1837 just as Queen Victoria came to the throne, but then had its proofs to correct. Charles Darwin’s health suffered from the pressure. On 20 September 1837, he had palpitations of the heart. On doctor’s advice that a month of recuperation was needed, Charles Darwin went to Shrewsbury then on to visit his Wedgwood relatives at Maer Hall, but found them too eager for tales of his travels to give him much rest. His charming, intelligent, and cultured cousin Emma Wedgwood, nine months older than Darwin, was nursing his invalid aunt. His uncle Jos pointed out an area of ground where cinders had disappeared under loam and suggested that this might have been the work of earthworms. This inspired a talk which Charles Darwin gave to the Geological Society on 1 November, the first demonstration of the role of earthworms in soil formation. William Whewell pushed Charles Darwin to take on the duties of Secretary of the Geological Society. After first declining this extra work, he accepted the post in March 1838. Despite the grind of writing and editing the Beagle reports, remarkable progress was made on transmutation. Darwin took every opportunity to question expert naturalists and, unconventionally, people with practical experience such as farmers and pigeon fanciers. Over time his research drew on information from his relatives and children, the family butler, neighbours, colonists and former shipmates. Charles Darwin included mankind in his speculations from the outset, and on seeing an ape in the zoo on 28 March 1838 noted its child-like behaviour. The strain took its toll, and by June he was being laid up for days on end with stomach problems, headaches and heart symptoms. For the rest of his life, he was repeatedly incapacitated with episodes of stomach pains, vomiting, severe boils, palpitations, trembling and other symptoms, particularly during times of stress, such as when attending meetings or dealing with controversy over his theory. The cause of Darwin’s illness was unknown during his lifetime, and attempts at treatment had little success. Recent attempts at diagnosis have suggested Chagas disease caught from insect bites in South America, Ménière’s disease, or various psychological illnesses as possible causes, without any conclusive results. On 23 June 1838, Charles Darwin took a break from the pressure of work and went geologising in Scotland. He visited Glen Roy in glorious weather to see the parallel roads cut into the hillsides at three heights. Charles Darwin thought that these were marine raised beaches: they were later shown to have been shorelines of a proglacial lake.

Fully recuperated, Charles Darwin returned to Shrewsbury in July. Used to jotting down daily notes on animal breeding, he scrawled rambling thoughts about career and prospects on two scraps of paper, one with columns headed Marry and Not Marry. Advantages included constant companion and a friend in old age ... better than a dog anyhow, against points such as less money for books and terrible loss of time. Having decided in favour, Charles Darwin discussed it with his father, then went to visit Emma on 29 July 1838. He did not get around to proposing, but against his father’s advice he mentioned his ideas on transmutation. Continuing his research in London, Darwin’s wide reading now included for amusement the 6th edition of Malthus’s An Essay on the Principle of Population which calculates from the birth rate that human population could double every 25 years, but in practice growth is kept in check by death, disease, wars and famine. Charles Darwin was well prepared to see at once that this also applied to de Candolle’s warring of the species of plants and the struggle for existence among wildlife, explaining how numbers of a species kept roughly stable. As species always breed beyond available resources, favourable variations would make organisms better at surviving and passing the variations on to their offspring, while unfavourable variations would be lost. This would result in the formation of new species. On 28 September 1838, Charles Darwin noted this insight, describing it as a kind of wedging, forcing adapted structures into gaps in the economy of nature as weaker structures were thrust out. Charles Darwin now had a theory by which to work, and over the following months compared farmers picking the best breeding stock to a Malthusian Nature selecting from variants thrown up by chance so that every part of every newly acquired structure is fully practised and perfected, and thought this analogy the most beautiful part of my theory.

On 11 November, Charles Darwin returned to Maer and proposed to Emma, once more telling her his ideas. She accepted, then in exchanges of loving letters she showed how she valued his openness, but her upbringing as a very devout Anglican led her to express fears that his lapses of faith could endanger her hopes to meet in the afterlife. While Charles Darwin was house-hunting in London, bouts of illness continued and Emma wrote urging him to get some rest, almost prophetically remarking So don’t be ill any more my dear Charley till I can be with you to nurse you. Charles Darwin found what they called Macaw Cottage (because of its gaudy interiors) in Gower Street, then moved his museum in over Christmas. The marriage was arranged for 24 January 1839, but the Wedgwoods set the date back. On the 24th, Darwin was honoured by being elected as Fellow of the Royal Society. On 29 January 1839, Charles Darwin and Emma Wedgwood were married at Maer in an Anglican ceremony arranged to suit the Unitarians, then immediately caught the train to London and their new home.

Charles Darwin now had the framework of his theory of natural selection by which to work, as his prime hobby. Charles Darwin research subsequently included animal husbandry and extensive experiments with plants, investigating many detailed ideas and finding evidence that species were not fixed to convince sceptical naturalists. For more than a decade this work was in the background to his main occupation, publication of the scientific results of the Beagle voyage. When FitzRoy’s Narrative was published in May 1839, Charles Darwin’s Journal and Remarks (The Voyage of the Beagle) as the third volume was such a success that later that year it was published on its own. Early in 1842, Charles Darwin sent a letter about his ideas to Lyell, who was dismayed that his ally now denied seeing a beginning to each crop of species. In May, Darwin’s book on coral reefs was published after more than three years of work, and he then wrote a pencil sketch of his theory. To escape the pressures of London, the family moved to rural Down House in November. On 11 January 1844 Darwin mentioned his theorising to the botanist Joseph Dalton Hooker, writing with melodramatic humour it is like confessing a murder. To Charles Darwin relief, Hooker replied There may in my opinion have been a series of productions on different spots, and also a gradual change of species. I shall be delighted to hear how you think that this change may have taken place, as no presently conceived opinions satisfy me on the subject.

By July, Darwin had expanded his sketch into a 230-page Essay, to be expanded with his research results if he died prematurely. Charles Darwin was shocked in November to find many of his arguments anticipated in the anonymously published Vestiges of the Natural History of Creation, though it lacked any convincing explanation for transmutation. The book was amateurish and he scorned its geology and anatomy, but as a best-seller it widened middle-class interest in transmutation, paving the way for Darwin as well as reminding him of the need to counter all arguments. Darwin completed his third geological book in 1846, and turned in relief to dissecting and classifying the barnacles he had collected, using his new ideas of common descent, and the anatomy he had learnt as Grant’s student. In 1847, Hooker read the Essay and sent notes that provided Darwin with the calm critical feedback that he needed, but would not commit himself and questioned Darwin’s opposition to continuing acts of Creation. In an attempt to improve his chronic ill health, Darwin went in 1849 to Dr. James Gully’s Malvern spa and was surprised to find some benefit from hydrotherapy. Then in 1851 his treasured daughter Annie fell ill, reawakening his fears that his illness might be hereditary. After a long series of crises, she died and Darwin’s faith in Christianity dwindled away. In eight years of work on barnacles (Cirripedia), Darwin found homologies that supported his theory by showing that slightly changed body parts could serve different functions to meet new conditions. In 1853 it earned Charles Darwin the Royal Society’s Royal Medal, and it made his reputation as a biologist. He resumed work on his theory of species in 1854, and in November realised that divergence in the character of descendants could be explained by them becoming adapted to diversified places in the economy of nature.

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By the start of 1856, Charles Darwin was investigating whether eggs and seeds could survive travel across seawater to spread species across oceans. Hooker increasingly doubted the traditional view that species were fixed, but their young friend Thomas Henry Huxley was firmly against evolution. Lyell was intrigued by Charles Darwin’s speculations without realising their extent. When Charles Darwin read a paper by Alfred Russel Wallace on the Introduction of species, he saw similarities with Darwin’s thoughts and urged him to publish to establish precedence. Though Darwin saw no threat, he began work on a short paper. Finding answers to difficult questions held him up repeatedly, and he expanded his plans to a big book on species titled Natural Selection. He continued his researches, obtaining information and specimens from naturalists worldwide including Wallace who was working in Borneo. In December 1857, Charles Darwin received a letter from Wallace asking if the book would examine human origins. Charles Darwin responded that he would avoid that subject, so surrounded with prejudices, while encouraging Wallace’s theorising and adding that I go much further than you. Charles Darwin’s book was half way when, on 18 June 1858, he received a paper from Wallace describing natural selection. Shocked that he had been forestalled, Darwin sent it on to Lyell, as requested, and, though Wallace had not asked for publication, offered to send it to any journal that Wallace chose. His family was in crisis with children in the village dying of scarlet fever, and he put matters in the hands of Lyell and Hooker. They agreed on a joint presentation at the Linnean Society on 1 July of On the Tendency of Species to form Varieties; and on the Perpetuation of Varieties and Species by Natural Means of Selection; however, Darwin’s baby son died of the scarlet fever and he was too distraught to attend. There was little immediate attention to this announcement of the theory; the president of the Linnean remarked in May 1859 that the year had not been marked by any revolutionary discoveries. Later, Darwin could only recall one review; Professor Haughton of Dublin claimed that all that was new in them was false, and what was true was old. Charles Darwin struggled for thirteen months to produce an abstract of his big book, suffering from ill health but getting constant encouragement from his scientific friends. Lyell arranged to have it published by John Murray. On the Origin of Species by Means of Natural Selection, or The Preservation of Favoured Races in the Struggle for Life (usually abbreviated to The Origin of Species) proved unexpectedly popular, with the entire stock of 1,250 copies oversubscribed when it went on sale to booksellers on 22 November 1859. In the book, Charles Darwin set out one long argument of detailed observations, inferences and consideration of anticipated objections. Charles Darwin only allusion to human evolution was the understatement that light will be thrown on the origin of man and his history. Charles Darwin theory is simply stated in the introduction: As many more individuals of each species are born than can possibly survive; and as, consequently, there is a frequently recurring struggle for existence, it follows that any being, if it vary however slightly in any manner profitable to itself, under the complex and sometimes varying conditions of life, will have a better chance of surviving, and thus be naturally selected. From the strong principle of inheritance, any selected variety will tend to propagate its new and modified form. Charles Darwin put a strong case for common descent, but avoided the then controversial term evolution, and at the end of the book concluded that; There is grandeur in this view of life, with its several powers, having been originally breathed into a few forms or into one; and that, whilst this planet has gone cycling on according to the fixed law of gravity, from so simple a beginning endless forms most beautiful and most wonderful have been, and are being, evolved.

There was wide public interest in Charles Darwin’s book and a controversy which he monitored closely, keeping press cuttings of reviews, articles, satires, parodies and caricatures. Critical reviewers were quick to pick out the unstated implications of men from monkeys, while amongst favourable responses Huxley’s reviews included swipes at Richard Owen, leader of the scientific establishment Huxley was trying to overthrow. Owen responded in April in a review which condemned the book. The Church of England scientific establishment, including Darwin’s old Cambridge tutors Sedgwick and Henslow, reacted against the book, though it was well received by a younger generation of professional naturalists. In 1860, the publication of Essays and Reviews by seven liberal Anglican theologians diverted clerical attention away from Darwin. An explanation of higher criticism and other heresies, it included the argument that miracles broke God’s laws, so belief in them was atheistic and praise for Mr Darwin’s masterly volume [supporting] the grand principle of the self-evolving powers of nature. The most famous confrontation took place at the public 1860 Oxford evolution debate during a meeting of the British Association for the Advancement of Science. Professor John William Draper delivered a long lecture about Darwin and social progress, then Samuel Wilberforce, the Bishop of Oxford, argued against Darwin. In the ensuing debate Joseph Hooker argued strongly for Darwin and Thomas Huxley established himself as Darwin’s bulldog the fiercest defender of evolutionary theory on the Victorian stage. Both sides came away feeling victorious, but Huxley went on to make much of his claim that on being asked by Wilberforce whether he was descended from monkeys on his grandfather’s side or his grandmother’s side, Huxley muttered: The Lord has delivered him into my hands and replied that he would rather be descended from an ape than from a cultivated man who used his gifts of culture and eloquence in the service of prejudice and falsehood.

Charles Darwin’s illness kept him away from the public debates, though he read eagerly about them and mustered support through correspondence. Asa Gray persuaded a publisher in the United States to pay royalties, and Charles Darwin imported and distributed Gray’s pamphlet Natural Selection is not inconsistent with Natural Theology. In Britain, friends including Hooker and Lyell took part in the scientific debates which Huxley pugnaciously led to overturn the dominance of clergymen and aristocratic amateurs under Owen in favour of a new generation of professional scientists. Owen made the mistake of (wrongly) claiming certain anatomical differences between ape and human brains, and accusing Huxley of advocating Ape Origin of Man. Huxley gladly did just that, and his campaign over two years was devastatingly successful in ousting Owen and the old guard. Charles Darwin’s friends formed The X Club and helped to gain him the honour of the Royal Society’s Copley Medal in 1864. Broader public interest had already been stimulated by Vestiges, and the Origin of Species was translated into many languages and went through numerous reprints, becoming a staple scientific text accessible both to a newly curious middle class and to working men who flocked to Huxley’s lectures. Darwin’s theory also resonated with various movements at the time and became a key fixture of popular culture.

Despite repeated bouts of illness during the last twenty-two years of his life, Charles Darwin pressed on with his work. Charles Darwin had published an abstract of his theory, but more controversial aspects of his big book were still incomplete, including explicit evidence of humankind’s descent from earlier animals, and exploration of possible causes underlying the development of society and of human mental abilities. Charles Darwin had yet to explain features with no obvious utility other than decorative beauty. His experiments, research and writing continued. When Charles Darwin’s daughter fell ill, he set aside his experiments with seedlings and domestic animals to accompany her to a seaside resort where he became interested in wild orchids. This developed into an innovative study of how their beautiful flowers served to control insect pollination and ensure cross fertilisation. As with the barnacles, homologous parts served different functions in different species. Back at home, Charles Darwin lay on his sickbed in a room filled with experiments on climbing plants. A reverent Ernst Haeckel who had spread a version of Darwinismus in Germany visited him. Wallace remained supportive, though he increasingly turned to Spiritualism. Variation of Plants and Animals Under Domestication, the first part of Darwin’s planned big book (expanding on his abstract published as The Origin of Species), grew to two huge volumes, forcing him to leave out human evolution and sexual selection, and sold briskly despite its size. A further book of evidence, dealing with natural selection in the same style, was largely written, but remained unpublished until transcribed in 1975.

The question of human evolution had been taken up by his supporters (and detractors) shortly after the publication of The Origin of Species, but Darwin’s own contribution to the subject came more than ten years later with the two-volume The Descent of Man, and Selection in Relation to Sex published in 1871. In the second volume, Darwin introduced in full his concept of sexual selection to explain the evolution of human culture, the differences between the human sexes, and the differentiation of human races, as well as the beautiful (and seemingly non-adaptive) plumage of birds. A year later Darwin published his last major work, The Expression of the Emotions in Man and Animals, which focused on the evolution of human psychology and its continuity with the behaviour of animals. He developed his ideas that the human mind and cultures were developed by natural and sexual selection, an approach which has been revived in the last three decades with the emergence of evolutionary psychology. As he concluded in Descent of Man, Darwin felt that, despite all of humankind’s noble qualities and exalted powers: Man still bears in his bodily frame the indelible stamp of his lowly origin. His evolution-related experiments and investigations culminated in books on the movement of climbing plants, insectivorous plants, the effects of cross and self fertilisation of plants, different forms of flowers on plants of the same species, and The Power of Movement in Plants. In his last book, he returned to the effect earthworms have on soil formation. He died in Downe, Kent, England, on 19 April 1882. He had expected to be buried in St Mary’s churchyard at Downe, but at the request of Darwin’s colleagues, William Spottiswoode (President of the Royal Society) arranged for Darwin to be given a state funeral and buried in Westminster Abbey, close to John Herschel and Isaac Newton.

The Darwins had ten children: two died in infancy, and Annie's death at the age of ten had a devastating effect on her parents. Charles was a devoted father and uncommonly attentive to his children. Whenever they fell ill he feared that they might have inherited weaknesses from inbreeding due to the close family ties he shared with his wife and cousin, Emma Wedgwood. He examined this topic in his writings, contrasting it with the advantages of crossing amongst many organisms. Despite his fears, most of the surviving children went on to have distinguished careers as notable members of the prominent Darwin-Wedgwood family. Of his surviving children, George, Francis and Horace became Fellows of the Royal Society, distinguished as astronomer, botanist and civil engineer, respectively. His son Leonard, on the other hand, went on to be a soldier, politician, economist, eugenicist and mentor of the statistician and evolutionary biologist Ronald Fisher.

Following Charles Darwin’s publication of the Origin, his cousin, Francis Galton, applied the concepts to human society, starting in 1865 with ideas to promote hereditary improvement which he elaborated at length in 1869. In The Descent of Man Charles Darwin agreed that Galton had demonstrated the probability that talent and genius in humans was inherited, but dismissed the social changes Galton proposed as too utopian. Neither Galton nor Charles Darwin supported government intervention and thought that, at most, heredity should be taken into consideration by people seeking potential mates. In 1883, after Darwin’s death, Galton began calling his social philosophy Eugenics.

During Charles Darwin’s lifetime, many species and geographical features were given his name. An expanse of water adjoining the Beagle Channel was named Darwin Sound by Robert FitzRoy after Darwin’s prompt action, along with two or three of the men, saved them from being marooned on a nearby shore when a collapsing glacier caused a large wave that would have swept away their boats, and the nearby Mount Darwin in the Andes was named in celebration of Darwin’s 25th birthday. When the Beagle was surveying Australia in 1839, Darwin’s friend John Lort Stokes sighted a natural harbour which the ship’s captain Wickham named Port Darwin. The settlement of Palmerston founded there in 1869 was officially renamed Darwin in 1911. It became the capital city of Australia’s Northern Territory, which also boasts Charles Darwin University and Charles Darwin National Park. Darwin College, Cambridge, founded in 1964, was named in honour of the Darwin family, partially because they owned some of the land it was on. The 14 species of finches he collected in the Galápagos Islands are affectionately named Darwin’s finches in honour of his legacy. Charles Darwin came fourth in the 100 Greatest Britons poll sponsored by the BBC and voted for by the public. In 2000 Charles Darwin’s image appeared on the Bank of England ten pound note, replacing Charles Dickens. His impressive, luxuriant beard (which was reportedly difficult to forge) was said to be a contributory factor to the bank’s choice. As a humorous celebration of evolution, the annual Darwin Award is bestowed on individuals who improve our gene pool by removing themselves from it. Numerous biographies of Darwin have been written, and the 1980 biographical novel The Origin by Irving Stone gives a closely researched fictional account of Darwin’s life from the age of 22 onwards.

Charles Darwin was a prolific author, and even without publication of his works on evolution would have had a considerable reputation as the author of The Voyage of the Beagle, as a geologist who had published extensively on South America and had solved the puzzle of the formation of coral atolls, and as a biologist who had published the definitive work on barnacles. While The Origin of Species dominates perceptions of his work, The Descent of Man, and Selection in Relation to Sex and The Expression of Emotions in Man and Animals had considerable impact, and his books on plants including The Power of Movement in Plants were innovative studies of great importance, as was his final work on The Formation of Vegetable Mould Through the Action of Worms.

Quotes by Charles Darwin:

"In the struggle for survival, the fittest win out at the expense of their rivals because they succeed in adapting themselves best to their environment."....Quotes by Charles Darwin.

"I have called this principle, by which each slight variation, if useful, is preserved, by the term Natural Selection"....Quotes by Charles Darwin.

"We will now discuss in a little more detail the Struggle for Existence."....Quotes by Charles Darwin.

"I love fools’ experiments. I am always making them."....Quote by Charles Darwin.

The Biography of Charles Darwin

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